I don't know about your meters, but all three of my Pentax digital spot meters (modified and unmodified) have fairly significant flare that will skew the meter reading for a dark object when there are brighter things around. This could easily be the cause of the underexposure the OP has. I have to be careful of this in the field, often compensating for possible flare by adding a bit of exposure. A lens hood on the meter helps a bit, but doesn't solve the problem entirely. FWIW, my modified meter tends to flare more than the unmodified (maybe all the filters added scatter more light?).
There are a number of other things that might influence the test results as well: faster shutter speeds than expected, inaccurate densitometer readings, development times that are on the short side, developer exhaustion or formulation not as expected (e.g., I had a batch of PMK that was inexplicably weak for no apparent reason), etc., etc.
My advice would be to err a bit on the overexposure side, go out and make some exposures and see how they print, paying special attentions to the shadow detail desired. Basing exposure on a Zone III or IV placement is a lot different that metering for Zone I and eliminates a lot of the flare problem.
I used to do a lot of film-speed testing, but these days just rate my film 2/3 stop slower than box speed to start, keep good notes and modify as I go. More important is finding a good "normal" development time that allows the maximum contrast control in the darkroom. Again, keeping notes and adjusting as I go works well for me.
Best,
Doremus