NOT a developer filling issue no hard edged well defined LINE would demark the transition of two fills!
Not an issue about curtain travel being 'inconsistent', else one would see multiple zones of higher and lower density as the slit moved inconsistently (slower in some places, faster in others) across the film plane.
I would say that the first curtain moves across first, but the second curtain gets initally delayed in its progression across the film plane...causing the first half of the area to get higher density than the second half