Kirk; Using the resolution charts that I have, the positive and negative exposures should match to determine the effect properly.
Rob; Excellent description of MTF. I'm not sure that what the chart shows is of line pairs though. The chart I have attached is more typical of line pairs. It is also more commonly called a definition chart as the definition of the film or print is determined by the last resolvable set of pairs. There are usually 4 lines or a set of "pairs" in this case.
The chart shown in the article is a typical MTF chart which is read at multiple densities to give resolution as a function of exposure. Reading two lines of unequal widths is, in this case, called a line pair which is a misnomer. So, in some cases it depends on the math used for analysis, the definition used, and not the chart which is clearly NOT line pairs.
The chart I have posted here shows pairs, but is never (AFAIK) used for MTF plots.
PE