To clarify, it looks to me like particles that fell from the bellows onto the surface of the film before you took the picture. There’s sticks and specks that look like they could be shellac crumbles. When you walkabout with a folding camera, sometimes you will open the bellows, setup and decide against taking a picture, close and walk on... Or you might walk around with camera open and on the tripod, jabbing the tripod down and maybe repositioning. All this action dislodges dust and specks from inside the camera body and bellows until you get many specks of clear spots on the negative.
If you wind just before taking the picture it won’t happen, though I can’t bear to do that. Next best plan is to shoot two or more shots in a short time. Definitely clean and vacuum bellows before going out. Replace the bellows if you have to