Maybe a clue, or maybe just a red herring, but the closest match to this I have found so far is the following, taken with a different camera on Velvia 100 (expired) and processed commercially:
View attachment 289230
This frame was also underexposed, a bit worse than the b&w above. The vertical bars aligned with the sprocket hole pattern are clearly here, especially in the blue channel, although I do not see them on the upper margin. A separate mystery, or part of the puzzle?