Here is my perspective regarding features and functionality.
The FE has a bright, uncluttered, logical, unobstructed viewfinder, with full information, including aperture and shutter speed (both selected and suggested). I have used dozens of camera bodies--Nikon, Pentax, Minolta, and Canon--and the FE has, in my opinion, the best, most useful finder. I do not have to press a button to observe this information, either.
The shutter in the FE is also quieter than what I had observed in the R4. See the review link I posted in my previous message, as that user's findings are similar to mine.
Further, the FE has a better accessory set, especially in regard to the motor drive. I paid $16 for my MD-11 (an EX grade unit from KEH, not a beat up "bargain" from an eBay seller), and it is perhaps the best drive I have used. It is compact and has a built in grip (not an optional grip). It features all-metal construction (I once easily cracked the plastic top piece on the more expensive MD-12), a 3.5/sec. top speed, a remote socket, and a vertical release attachment feature. Better yet, the attachment of either an MD-11 or MD-12 makes the FE a "one switch" camera--the motor's power switch runs the whole rig. The only thing the MD-11 lacks (which the MD-12 has) is an automatic turn-off circuit. I have yet to have difficulty with this, as I just remind myself to turn off the unit when done. I did not have to pay $200 for the motor, or search high and low for one in decent shape, either.
With regard to lenses, the FE offers me the option of either non-AI or AI types. Thus, I may choose from numerous of classic and newer lenses, depending upon what I am trying to achieve (single-coated vs. multicoated, built-in hood vs. separate hood, etc.).
In short, I have yet to find a better system overall, although my Minolta XG-M/MD-1 setup is quite excellent, too. The XG-M has a nicer shutter release--nice and soft--and the vertical release on the MD-1 is simply outstanding (soft and well positioned) for shots in the vertical format.