Yes, they are the rollers, and thanks for posting the photos.
If you have a micro cloth, like the one used to clean reading glasses, then you could rub along the rollers, then rotate the roller, a 1/4 rotation, clean again along the roller and repeat until you have both rollers cleaned.
If there are any stubborn spots (you will probably feel them as you clean even if you don't see them) then use a drop of methylated spirits/denatured alcohol and it should do the trick.
At least it will eliminate the rollers as a cause if the problem persists.
Let us know how you get on.