There are five cameras where the difference between AI and AIS matters, as listed here:
http://www.throughthefmount.com/articles_back_difference_ai_ais.html
I guess that article reinforces my point that there are only 5 cameras that can tell the difference between an AI and AI-S lens. Of those the F4 simply ignores most of the features of an AI-S lens, namely it ignores the linear aperture, as it will not let you control the aperture without a cpu. The FA gets around it by using a closed circuit meter at exposure time. That just leaves the FG, F-301, F-501.
As for the D700, it definitely can control the aperture on CPU-based lenses, which it does for my 60/2.8 G Micro Nikkor.
I said the D700 can't control the aperture of a non-CPU based lens, not the other way around. If it can it would be nice to know. According to the article you posted I think all DSLRs are the same.
Edit,
The article suggests there are only two cameras the FA and F-501 where it makes a difference...
https://www.graysofwestminster.co.uk/glossary/ai_ais_difference.php
The only two camera models actually make full use of the Ai-S lens mount: the Nikon FA and F-501, both of which have been out of production for many years. The F-301 and F4 will detect that an Ai-S type lens has been mounted but both these cameras operate with Ai lenses in an identical way. All other Nikon cameras make no differentiation between the Ai and Ai-S mount.
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