Most plastic reels I've seen (but not all) use the system where one rotates against the other, while little ball bearings hold the film after it advances. I've never liked these because I always find the film jams. I'm wondering if the marks aren't torsion stretch marks in the emulsion caused by the twisting motion of loading the film.
I'd try a metal reel and see if it presents again.
I vote for a tiny crack or other light leak in your developing tank. It repeats because the reels move or rotate slightly during agitation.
Peter Gomena
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