My hypothesis:
The film end closest to the center was being submerged in the developer more often than the outer portions of the film, causing a gradation of development.
This would be true only if you loaded the reels all the way to the inside of the reel, i.e. all the way to the center core, leaving the empty space on the outer part of the reel - which is NOT how you load the film if you want to use rotation. If you use "normal" way of "hand" developing in a tank, it doesn't matter. But, for rotation, the film must not be threaded all the way to the core. Or, it may - but then you must fill the tank/drum almost completely - according to instructions, which can be found on the web, e.g.
HERE.
You'll see that the quantity of chemicals if you use ONE 120 film per reel is lower than if you want to use TWO films per reel.
Some (newer) Jobo reels have a red tab, with instructions that you load the 120 film only until you reach the red tab. The way to load it is to "thread" the film in only until you reach the end of the film - do NOT thread it any more - i.e. closer to the center. The red tab is used when you load TWO 120 films on the same reel, preventing overlapping of the negatives.
Jobo instructions say the following:
"Continue walking the film onto the reel until the end is fully into the reel."
(by "walking" I guess they mean "threading", or whatever you want to call the method by which you insert the roll onto the reel).
Denis