I recently printed a few contact sheets (from 4x5" sheet film) and then noticed what appears to be fingerprints on one of the negatives (see image attached, to the left and right and just below the horizon). It looks like the marks are in the emulsion. I am a bit puzzled since the marks have built extra density compared to the background; I would expect greasy fingerprints applied pre-development to result in developer not reaching the emulsion end hence less density. Can greasy fingerprints applied post-development somehow build extra density to the emulsion? Most importantly, is there some way to remove the marks? I tried to gently rub the negative with a cotton ball soaked in IPA, without success...