Only on long exposures? Have you had any printed by a lab? If when printed they still appear then it is the camera, lens or filter if one was on the lens or processing. If not then it is something with the scanner, computer or handling after processing.
http://www.jeffreyglasser.com/
Was it humid when you took the photo, or is that a bad question since you appear to live in England? Lol.
My guess is the film is getting "sticky" when exposed to humid air for an extended time and is reacting to the backing paper, or sticking slightly to the backing paper. Backing paper ain't made like it used to be.... If that is the case then the only thing I can think of is to do a longer pre wet. Not sure if that will help though.
Or when you look at the emulsion side in oblique light? perhaps moisture condensed on the film, then the film was advanced trapping the moisture between layers of film on the takeup spool. See if you can isolate it to the emulsion side. The exposures are obviously minutes long, a lot can happen inside the camera under the right (or wrong) circumstances.Can you see any sign of the spots when you look at the film itself under high magnification?
Mamiya RZ67 pro 2, f32, 16 minutes exposure with dark sky.
My first thought was they could be star trails from the long exposure, specifically more distant stars moving into and out of the view. This is common (and normal).
You would not get many trails of different colours until you go up to about 4 hours in exposure, and up to 6 hours or more trails that encircle the entire image space.
Star trails are best at shallow apertures e.g. f5.6 to f.8. I am unsure why you went to f32.
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