The blades have pins, and if the pins dislodge from any of the blades, the opening of the aperture will become irregular. There's two pins for each blade, one is stationary, the other follows along the cam slot, and this is the one that usually dislodges. One of mine has a pin missing, it's another repair I must do, besides making new shutter blades, to completely fix the shutter. Quite often, a blade with a missing pin will still open and close due to friction alone, or there may be a burr around the hole of the missing pin, and that can be enough to "catch" the cam, and act like the pin is still there. But of course it's not, but the affected blade will be a little out of alignment, causing the aperture opening to be just that little bit out of round.